Uusi opinto-opas (sisältäen myös opetusohjelmat) lukuvuodelle 2018-2019 sijaitsee osoitteessa https://opas.peppi.utu.fi . Tältä sivustolta löytyvät enää vanhat opinto-oppaat ja opetusohjelmat.

The new study guide (incl. teaching schedules) for academic year 2018-2019 can be found at https://studyguide.utu.fi. This site contains only previous years' guides.

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Arkistoitu opetusohjelma 2016–2017
Selaat vanhentunutta opetusohjelmaa. Voimassa olevan opetusohjelman löydät täältä.
BIOL4321 Advanced evolutionary biology 1 5 ECTS
Period I Period II Period III Period IV
Language of instruction
Type or level of studies
Advanced studies
Course unit descriptions in the curriculum
Department of Biology

General description

The course will begin with an overview of the genetic theory of natural selection and discuss ways to detect natural selection at the molecular level. Will then look at the genetic architecture of phenotypic traits and get an overview of quantitative trait evolution, getting also into polygenic traits. At this stage will get into the evolution at different molecular levels: genomic, transcriptomic, proteomic and on current models to detect selection at each level. Will then change direction and discuss the paradox of sex and about sexual selection. Will proceed with the concept of species and take the opportunity to get into comparative genomics, and also into theories of the effects of hybridization and genomic introgression. Will have then a look at the evolution of genes and of genomes before concluding with concepts about the evolution of cooperation, kin selection, group selection, and about macroevolution. This is a general description of the course, and does not include content of exercises assays and practical training that may be part of the training of the students.

Teacher responsible

Spiros Papakostas


Mon 3-Oct-2016 - 17-Oct-2016 weekly at 10-12, Natura bld, 2nd floor, room 215 (Seminar Room)
Wed 5-Oct-2016 - 19-Oct-2016 weekly at 12-14, Natura bld, 2nd floor, room 215 (Seminar Room)
Mon 24-Oct-2016 - 12-Dec-2016 weekly at 10-12, Natura bld, 2nd floor, room 251 (ecology corridor)
Wed 26-Oct-2016 - 14-Dec-2016 weekly at 12-14, Natura bld, 2nd floor, room 251 (ecology corridor)

Further information

What prior knowledge the students should have

Students should have knowledge of the fundamental principles of modern evolutionary synthesis such as the mendelian laws of inheritance, sources of genetic variation (standing genetic variation, mutation, introgression). Also, students should be well-aware of what is natural selection and genetic drift, and should be knowledgeable of basic population genetics (e.g. about the HW principle or what is standing genetic variation, hard and soft sweeps, what is genetic hitchhiking etc.). Knowledge about the concepts of molecular clock and phylogenetics is also desirable but not necessary to follow the course. This course is aimed to be a continuation of the Evolutionary Biology course taught to undergraduates, so local students should have followed that. Basic knowledge of R is a plus but not necessary to follow the course