Uusi opinto-opas (sisältäen myös opetusohjelmat) lukuvuodelle 2018-2019 sijaitsee osoitteessa https://opas.peppi.utu.fi . Tältä sivustolta löytyvät enää vanhat opinto-oppaat ja opetusohjelmat.

The new study guide (incl. teaching schedules) for academic year 2018-2019 can be found at https://studyguide.utu.fi. This site contains only previous years' guides.

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Arkistoitu opetussuunnitelma 2013–2014
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BIOI4390 Major Transitions in Evolution 3 op
Organised by
Person in charge
Csaba Ortutay
Preceding studies
Elementary knowledge of molecular machinery of inheritance (base pairing, transcription, translation). A concept on mechanism of selection and basic knowledge about natural history of life.

Learning outcomes

Overview of most important events in molecular evolution of life.


Formation of life; Forming of replicators; Forming of chromosomes; Switch from RNA to DNA; The Prokaryotes - Eukaryotes transition; Origin of sexuality; Multicellular organisms; Formation of colonies, societies and the case of languages.

Teaching methods

Teaching method Contact Online
Lectures 20 h 0 h

Teaching language


Modes of study

Option 1
Available for:
  • Degree Programme Students
  • Other Students
  • Doctoral Students
  • Exchange Students
Oral exam
  • In English
Oral exam

Final examination on the lectures and selected parts of the literature.


Numeric 0-5.

Recommended year of study

2. year autumn

Study materials

Maynard Smith and Szathmáry (1995) The Major Transitions in Evolution.

Belongs to following study modules

Tulevaisuuden teknologioiden laitos
Tulevaisuuden teknologioiden laitos
Tulevaisuuden teknologioiden laitos
Archived Teaching Schedule. Please refer to current Teaching Shedule.
Implementation details are unavailable.
Tulevaisuuden teknologioiden laitos