Uusi opinto-opas (sisältäen myös opetusohjelmat) lukuvuodelle 2018-2019 sijaitsee osoitteessa https://opas.peppi.utu.fi . Tältä sivustolta löytyvät enää vanhat opinto-oppaat ja opetusohjelmat.

The new study guide (incl. teaching schedules) for academic year 2018-2019 can be found at https://studyguide.utu.fi. This site contains only previous years' guides.

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Arkistoitu opetussuunnitelma 2016–2018
Selaamasi opetussuunnitelma ei ole enää voimassa. Tarkista tiedot voimassa olevasta opetussuunnitelmasta.
BIOL4321 Advanced evolutionary biology 1 5 op
Organised by
Person in charge
The information is in the Teaching Schedules.
Preceding studies
Recommended prior courses: EKOL3159 Introduction to R, 4 op, BIOL2121 Evoluutiobiologia, 5 op or equivalent knowledge.

Learning outcomes

Students are familiar with the evolutionary concepts taught in the course issues.


Themes covered include (but are not restricted to) phenotypic and genetic variation: origin and maintenance, Natural selection and adaptation, Evolution of genes and genomes, evolutionary genomics, advanced evolutionary analyses using R and other programs, societal applications of evolutionary biology.

Teaching methods

Teaching method Contact Online

Lectures, student presentations and discussions on the presented topics. The course will (loosely) follow the text book Evolution, by Futuyma.

Teaching language


Modes of study

Oral presentation, active participation in discussions, and study journal/learning diary.


Numeric 0-5.

Recommended year of study

4. year autumn
5. year autumn

Further information

This course will be arranged every second autumn (even years). Sign up for the course in nettiopsu.utu.fi. Course will be arranged, if at least 8 students register for the course.

Belongs to following study modules

Biologian laitos
Archived Teaching Schedule. Please refer to current Teaching Shedule.
Implementation details are unavailable.
Biologian laitos